Maryland Darter

Etheostoma sellare

Percidae · freshwater water fish

5 minute read · Species guide

Maryland Darter field guide illustration

About Maryland Darter

The Maryland darter (Etheostoma sellare) is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is considered one of the rarest freshwater fish species in the world, due in part to its extremely limited geographic range and difficulty of detection. The last sighting of one was in 1988. The Maryland darter is named after the only state in which it is known to occur. The species was long known only by two specimens until being "re-discovered" in 1962. From 1965 into the 1980s, the species was believed to have been confined to a single riffle in Deer Creek. Possible explanations for the decline of the species center around widespread habitat degradation and reduction in water quality resulting from increasing rates of urbanization within the watershed. While the IUCN has declared the species extinct, the United States Fish and Wildlife Service has not, and keeps it on the Endangered Species List.

History

The Maryland darter was originally discovered in 1912 in Swan Creek, a tributary of the larger Susquehanna River drainage basin located near Havre de Grace, Maryland. The two biologists responsible for the discovery, Lewis Radcliffe and William W. Welsh, promptly published a description of the newly discovered species in 1913. Despite initial detection of the species in Swan Creek, it was never found there again. However, in 1962 the Maryland darter was "re-discovered" after an adult female specimen was collected in nearby Gasheys Run, a tributary of Swan Creek. Perhaps the most significant detection of the Maryland darter occurred in 1965, when over 70 individuals were collected from Deer Creek. Additional sightings of the species in Deer Creek occurred irregularly between 1974 and 1988, where between 1 and 10 individuals were detected by seining sampling or snorkeling. However, the last significant sighting of the Maryland darter occurred in 1988. It has not been seen since, and is believed to be extinct. This species is the only member of the subgenus Mooreichthys.

Characteristics

The Maryland darter is a member of the genus Etheostoma and a member of the darter subfamily of fishes. A relative of the yellow perch (Perca flavescens) and walleye (Sander vitreus), the Maryland darter is a bottom-dwelling fish. Ideal habitat consists of highly oxygenated, swiftly flowing portions of streams containing moderate amounts of vegetation and characterized by rubble and gravel-like substrate. Additionally, the species favors sloped rock riffles and crevasses which provide ample shelter from predators. The Maryland darter is a relatively smaller darter species with a maximum size of just under three inches. The species favors small invertebrates, frequently foraging on small snails, caddis fly larvae and mayfly and stonefly nymphs. Spawning presumably occurs in late April as is characteristic of most other darter species, however the Maryland darter has never been observed during reproduction.

Species status

The Maryland darter is listed as endangered by the United States Fish and Wildlife Service. The United States Fish and Wildlife Service has determined that due to difficulty of detection, in spite of infrequent observations since initial discovery in 1912, that there is not sufficient evidence to declare the species extinct. However, other organizations such as the IUCN has listed the Maryland darter as extinct on the IUCN list of threatened species. The state of Maryland has the species listed as SH, indicating that, historically, the species occurred in the state but is believed to be extirpated, having not been observed for a period of over 20 years.

The Maryland Darter is seemingly becoming extinct due to the high quantities of pollution from agricultural and development runoff. We can determine that sediment from erosion and runoff reduced water quality and may have smothered darter eggs. Erratic water levels created sedimentation problems; stranding fish in pools. These pools could become too warm for them to survive, which increases predation by other fish.

Recovery efforts

The Maryland Darter Recovery Plan is a plan implemented by the Wildlife and Heritage Service of the Maryland Department of Natural Resources, devised in cooperation with the United States Fish and Wildlife Service. The intent of the plan is to implement widespread detection efforts using a variety of techniques. including electro-fishing equipment and electro trawls, in an effort to learn more about this rare species. The plan also recommends the protection of the Deer Creek watershed through the use of improved farming and forestry practices. In addition, strips of naturally vegetated lands are to be constructed adjacent to Deer Creek in an effort to insulate the watershed from harmful nearby agricultural runoff. The root systems and leaf cover established by these buffers would, in addition to preventing erosion, would help to absorb toxic chemicals and filter solid waste before it has a chance to enter the watershed. Additionally, surveys have been conducted by the Maryland Biological Stream Survey (MBSS) in 2001 through 2006. Six sites in Deer Creek were surveyed by MBSS as well as 22 additional sites in the greater Deer Creek watershed. One site in Gasheys Run, one site in the mainstem of Swan Creek, and four more sites within the greater Swan Creek Watershed were also sampled. Again, all of these surveys resulted in inconclusive evidence. Attempts to locate the Maryland darter remain largely unsuccessful.

Article text: Wikipedia contributors, Etheostoma sellare (source revision). Licensed under CC BY-SA 4.0. Retrieved 2026-10-11.

Diet and feeding

The FishTraits compilation records these diet categories for this species: algae or phytoplankton; invertebrates or larval fish. These are broad categories, not a claim that every individual consumes every example in a category.

Reported feeding positions include bottom feeding. The compilation combines published observations; diets can vary with age, locality, and available food.

Habitat preferences

Reported habitat associations include streams and small rivers, creeks.

The source records associations with gravel, cobble, vegetation. An association indicates reported use, rather than an exclusive requirement.

Reproduction and parental care

The compilation reports a representative maturity age of 1 years. Its maturity field summarizes mean, median, or modal estimates, usually for females; male estimates can be used when female data are unavailable. It is not a fixed age at which every fish first breeds.

Recorded spawning months are Apr, May for the source’s reporting region (MD). This is a regional calendar, not a range-wide schedule.

Size and recorded longevity

FishTraits reports a maximum total length of 7 centimetres. Some entries use an estimated asymptotic length; this value should not be treated as a certified record catch.

Recorded longevity in the compilation is 3 years. Wild observations were preferred, but captive records were accepted where wild records were unavailable, and unspecified records were assumed wild. The value is not a guaranteed wild lifespan or an aquarium life expectancy.

Range

Where it lives

Share of 14 GBIF observation records with a state location, across 1 states.
0%
100%
Share of 14 GBIF observation records with a state location, across 1 states.

Share of records by state

Habitat

Water and movement

Water
WoRMS records this species in fresh water.
Migration
Undetermined

Classification

Taxonomy

Kingdom
Animalia
Phylum
Chordata
Superclass
Actinopterygii
Class
Teleostei
Order
Perciformes
Family
Percidae
Genus
Etheostoma
Species
Etheostoma sellare
ITIS TSN
168361
WoRMS AphiaID
1525897
GBIF key
2382401

Common questions about Maryland Darter

What is the scientific name of Maryland Darter?
The scientific name is Etheostoma sellare, in the family Percidae. Classification follows ITIS.
Does Maryland Darter live in fresh water or salt water?
WoRMS records this species in fresh water.
Where has Maryland Darter been recorded in the United States?
GBIF has observations in 1 states, with the most records in Maryland. The map shows observation records, which do not establish the complete natural range.

Closest relatives

Other Etheostoma species

Same family

More Percidae

All 229 species in Percidae